Lone Tree Milsim captured the intense action of American Milsim's Revolt event, where the Coalition of Sovereign Territories (COST) and the United Federal States (UFS) clashed in a pivotal showdown at the now-desolate Desert Center, California. This former stronghold of the Pacific State Alliance (PSA), once a powerful third faction, became the battleground for a fight over remaining sensitive materials, prompting both COST and UFS to mobilize massive forces, as detailed in Lone Tree Milsim's trailer.